info prev up next book cdrom email home

Life Expectancy

An $l_x$ table is a tabulation of numbers which is used to calculate life expectancies.

$x$ $n_x$ $d_x$ $l_x$ $q_x$ $L_x$ $T_x$ $e_x$
0 1000 200 1.00 0.20 0.90 2.70 2.70
1 800 100 0.80 0.12 0.75 1.80 2.25
2 700 200 0.70 0.29 0.60 1.05 1.50
3 500 300 0.50 0.60 0.35 0.45 0.90
4 200 200 0.20 1.00 0.10 0.10 0.50
5 0 0 0.00 -- 0.00 0.00 --
$\sum$   1000 2.70        

$x$: Age category ($x=0$, 1, ..., $k$). These values can be in any convenient units, but must be chosen so that no observed lifespan extends past category $k-1$.

$n_x$: Census size, defined as the number of individuals in the study population who survive to the beginning of age category $x$. Therefore, $n_0=N$ (the total population size) and $n_k=0$.

$d_x$: $=n_x-n_{x+1}$; $\sum_{i=0}^k d_i=n_0$. Crude death rate, which measures the number of individuals who die within age category $x$.

$l_x$: $=n_x/n_0$. Survivorship, which measures the proportion of individuals who survive to the beginning of age category $x$.

$q_x$: $=d_x/n_x$; $q_{k-1}=1$. Proportional death rate, or ``risk,'' which measures the proportion of individuals surviving to the beginning of age category $x$ who die within that category.

$L_x$: $=(l_x+l_{x+1})/2$. Midpoint survivorship, which measures the proportion of individuals surviving to the midpoint of age category $x$. Note that the simple averaging formula must be replaced by a more complicated expression if survivorship is nonlinear within age categories. The sum $\sum_{i=0}^k L_x$ gives the total number of age categories lived by the entire study population.

$T_x$: $=T_{x-1}-L_{x-1}$; $T_0=\sum_{i=0}^k L_x$. Measures the total number of age categories left to be lived by all individuals who survive to the beginning of age category $x$.

$e_x$: $=T_x/l_x$; $e_{k-1}=1/2$. Life expectancy, which is the mean number of age categories remaining until death for individuals surviving to the beginning of age category $x$.


For all $x$, $e_{x+1}+1>e_x$. This means that the total expected lifespan increases monotonically. For instance, in the table above, the one-year-olds have an average age at death of $2.25+1=3.25$, compared to 2.70 for newborns. In effect, the age of death of older individuals is a distribution conditioned on the fact that they have survived to their present age.


It is common to study survivorship as a semilog plot of $l_x$ vs. $x$, known as a Survivorship Curve. A so-called $l_x m_x$ table can be used to calculate the mean generation time of a population. Two $l_x m_x$ tables are illustrated below.

\begin{figure}\begin{center}Population 1\end{center}\end{figure}

$x$ $l_x$ $m_x$ $l_x m_x$ $xl_xm_x$
0 1.00 0.00 0.00 0.00
1 0.70 0.50 0.35 0.35
2 0.50 1.50 0.75 1.50
3 0.20 0.00 0.00 0.00
4 0.00 0.00 0.00 0.00
      $R_0=1.10$ $\sum=1.85$


$\displaystyle T$ $\textstyle =$ $\displaystyle {\sum xl_xm_x\over\sum l_xm_x}={1.85\over 1.10} = 1.68$  
$\displaystyle r$ $\textstyle =$ $\displaystyle {\ln R_0\over T}={\ln 1.10\over 1.68} = 0.057.$  


\begin{figure}\begin{center}Population 2\end{center}\end{figure}

$x$ $l_x$ $m_x$ $l_x m_x$ $xl_xm_x$
0 1.00 0.00 0.00 0.00
1 0.70 0.00 0.00 0.00
2 0.50 2.00 1.00 2.00
3 0.20 0.50 0.10 0.30
4 0.00 0.00 0.00 0.00
      $R_0=1.10$ $\sum=2.30$


$\displaystyle T$ $\textstyle =$ $\displaystyle {\sum xl_xm_x\over\sum l_xm_x}={2.30\over 1.10} = 2.09$  
$\displaystyle r$ $\textstyle =$ $\displaystyle {\ln R_0\over T}={\ln 1.10\over 2.09} = 0.046.$  

$x$: Age category ($x=0$, 1, ..., $k$). These values can be in any convenient units, but must be chosen so that no observed lifespan extends past category $k-1$ (as in an $l_x$ table).

$l_x$: $=n_x/n_0$. Survivorship, which measures the proportion of individuals who survive to the beginning of age category $x$ (as in an $l_x$ table).

$m_x$: The average number of offspring produced by an individual in age category $x$ while in that age category. $\sum_{i=0}^k m_x$ therefore represents the average lifetime number of offspring produced by an individual of maximum lifespan.

$l_x m_x$: The average number of offspring produced by an individual within age category $x$ weighted by the probability of surviving to the beginning of that age category. $\sum_{i=0}^k l_xm_x$ therefore represents the average lifetime number of offspring produced by a member of the study population. It is called the net reproductive rate per generation and is often denoted $R_0$.

$xl_xm_x$: A column weighting the offspring counted in the previous column by their parents' age when they were born. Therefore, the ratio $T=\sum(xl_xm_x)/\sum(l_xm_x)$ is the mean generation time of the population.


The Malthusian Parameter $r$ measures the reproductive rate per unit time and can be calculated as $r=(\ln R_0)/T$. For an exponentially increasing population, the population size $N(t)$ at time $t$ is then given by

\begin{displaymath}
N(t)=N_0 e^{rt}.
\end{displaymath}

In the above two tables, the populations have identical reproductive rates of $R_0=1.10$. However, the shift toward later reproduction in population 2 increases the generation time, thus slowing the rate of Population Growth. Often, a slight delay of reproduction decreases Population Growth more strongly than does even a fairly large reduction in reproductive rate.

See also Gompertz Curve, Logistic Growth Curve, Makeham Curve, Malthusian Parameter, Population Growth, Survivorship Curve



info prev up next book cdrom email home

© 1996-9 Eric W. Weisstein
1999-05-25